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A U.S. Marine F-35 Lightning II from the Marine Fighter Attack Squadron (VMFA) 211 taxis down the Tyndall flightline during a 53rd Weapon Evaluation Group Combat Archer training mission Sept. 16, 2016. During this Weapon System Evaluation Program, VMFA 211 preformed the first operational firing of the F-35’s air-to-air missile weapons system. (U.S. Air Force photo by Senior Airman Solomon Cook/Released)

F-35 Lightning II has first operational air-to-air missile fire

A U.S. Marine F-35 Lightning II from the Marine Fighter Attack Squadron (VMFA) 211 taxis down the Tyndall flightline during a 53rd Weapon Evaluation Group Combat Archer training mission Sept. 16, 2016. During this Weapon System Evaluation Program, VMFA 211 preformed the first operational firing of the F-35’s air-to-air missile weapons system. (U.S. Air Force photo by Senior Airman Solomon Cook/Released)

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